Laboratory of Marine Symbiotic Biology, Kochi University
<Japanese >
Gyo Itani, Ph.D.

Associate Professor

Laboratory of Marine Symbiotic Biology
Faculty of Education, Kochi University
2-5-1 Akebono, Kochi 780-8520, Japan

tel: +81-88-844-8415
e-mail: itani@

1995. B.Sc. Kyoto University
1997. M.Sc. Kyoto University
2002. Ph.D. Kyoto University

Professional history
1997-2000. JSPS Research Fellow (DC1) at SMBL, Kyoto University
2003-2005. JSPS Research Fellow (PD) at CMES, Ehime University
2005-present. Associate Professor, Faculty of Education, Kochi University

Biology II
Organisms and their environments
Marine ecology I & II
Summer field courses (marine biology)
Biodiversity I & II (for postgraduates)

Ecology of thalassinidean shrimp
Ecology and taxonomy of bopyrid isopods
Ecology and taxonomy of symbiotic crabs (Sestrostoma)
Ecology of symbiotic clams (Peregrinamor and Cryptomya)

Atsushi Mino: Distributions of tidal flat brachyurans.
Yumi Henmi: Symbiotic ecology of gobies associated with shrimp burrows.
Jonel M. Corral: Parasitic effects of bopyrids on host brachyuran crabs.
Yuto Shiozaki: Symbiotic ecology of Sestrostoma crabs.
Sota Kirihara: Symbiotic ecology of goby associated with Alpheus shrimp.

Kazumi Oda: Burrow morphology of Upogebia sakaii.
Runa Murakami: Parasitic effects of Peregrinamor ohshimai on host shrimp.
Koki Nonaka: Symbiotic ecology of Stenalpheops anacanthus.

Upogebia major with Gyge ovalis and Peregrinamor ohshimai
Sestrostoma toriumii in the burrows of Upogebia major
Recent publications

Henmi, Y., Fujiwara, C., Kirihara, S., Okada, Y., and Itani, G. (2017) Burrow morphology of alpheid shrimps: Case study of Alpheus brevicristatus and a review of the genus. Zoological Science, in press.

Ijichi, M., Itani, G., and Ueda, H.(2017) Life cycle and precopulatory mate guarding of Goidelia japonica (Copepoda: Poecilostomatoida: Echiurophilidae) associated with the echiuran Urechis unicinctus. Plankton and Benthos Research, in press.

Henmi, Y., Eguchi, K., Inui, R., Nakajima, J., Onikura, N., and Itani, G. (2017) Field survey and resin casting of Gymnogobius macrognathos spawning nests in the Tatara River, Fukuoka Prefecture, Japan. Ichthyological Research, in press.

Henmi, Y., Okada, Y., and Itani, G. (2017): Field and laboratory quantification of alternative use of host burrows by the varunid crab Sestrostoma toriumii (Takeda, 1974) (Brachyura: Varunidae). Journal of Crustacean Biology, 37: 235-242.

Lutzen, J., Itani, G. Jespersen, A., Hong, J.-S., Rees, D., and Glenner, H. (2016) On a new species of parasitic barnacle (Crustacea: Rhizocephala), Sacculina shiinoi sp. nov., parasitizing Japanese mud shrimps Upogebia spp. (Decapoda: Thalassinidea: Upogebiidae), including a description of a novel morphological structure in the Rhizocephala.ĘZoological Science, 33: 204-212.

Tateda, Y., Sakaguchi, I. & Itani, G. (2015) Scope for growth of Mytilus galloprovincialis and Perna viridis as a thermal stress index in the coastal waters of Japan: Field studies at the Uranouchi inlet and Yokohama. Journal of Experimental Marine Biology and Ecology, 470: 55-63.

Henmi, Y. & Itani, G. (2014) Burrow utilization in the goby Eutaeniichthys gilli associated with the mud shrimp Upogebia yokoyai. Zoological Science, 31: 523-528.

Henmi, Y. & Itani, G. (2014) Laboratory quantification of burrow utilization by the symbiotic varunid crab Sestrostoma toriumii. Plankton and Benthos Research, 9: 203-206.

Henmi, Y., Iwata, Y. & Itani, G. (2014) Associations of the gobies Eutaeniichthys gilli and Gymnogobius scrobiculatus with burrows of the mud shrimp Upogebia yokoyai at low tide. Japanese Journal of Benthology, 69: 69-75. (in Japanese with English Abstract)

Itani, G., Yamada, C., Asama, H., Henmi, Y., Kume, H. and Chapman, J. W. (2014) Parasitic crustaceans and marine invasions: two case studies from Kuroshio region. Kuroshio Science, 8: 107-110.

Henmi, Y. & Itani, G. (2013) Records of the endangered crab Hexapinus latipes collected with Chaetopterus cautus in Uranouchi Inlet, Kochi, Japan. Nankiseibutu, 55: 115-117. (in Japanese with English Abstract)

Sato, A., Morinaga, J., Henmi, Y. & Itani, G. (2013) New record of the Japanese mud shrimp, Upogebia major, from Kochi Prefecture, Japan. Kuroshio Science, 6: 238-242. (in Japanese)

Ueda, I., Sakaguchi, I., Ogita, T., Yamada, C. & Itani, G. (2013) Thermal conditions of overwintering for tropical green mussel Perna viridis in Uranouchi Inlet, Tosa Bay, Shikoku, Japan in 2010. Sessile Organisms, 30: 29-36. (in Japanese)

Chapman, J. W., Dumbauld, B. R.,Itani, G. & Markham, J. C. (2012) An introduced Asian parasite threatens northeastern Pacific estuarine ecosystems. Biological Invasions,14: 1221-1236.

Itani, G., Yamada, C. & Watanabe, T. (2011) Utilization of the non-indigenous mussel, Mytilus galloprovincialis, by the native pea crab Arcotheres sinensis in 1930s revealed from the specimens in the museum of the Seto Marine Biological Laboratory, Kyoto University. Kuroshio Science, 4: 169-174. (in Japanese)

Reimer, J. D., Hirose, M., Nishisaka, T., Sinniger, F. & Itani, G. (2010): Epizoanthus spp. associations revealed using DNA markers: a case study from Kochi, Japan. Zoological Science, 27: 729-734.

Kinoshita, K., G. Itani & T. Uchino (2010) Burrow morphology and associated animals of the mud shrimp Upogebia yokoyai (Crustacea: Thalassinidea: Upogebiidae). Journal of the Marine Biological Association of the United Kingdom, 90: 947-952.

Hirota, F., G. Itani, M. Ikehara, H. Ueda & I. Kinoshita (2010) Planktonic foraminiferal assemblage in coastal water of Tosa Bay, southwestern Japan. Bulletin of Marine Sciences and Fisheries Kochi University, 26: 1-7. (in Japanese)

Yamada, C., T. Kakio, H. Kurita & G. Itani (2010) A comparison of the reproductive seasons of three mytilid species in Uranouchi Inlet, Kochi, Japan. Kuroshio Science, 3: 138-143. (in Japanese)

Yamada, C., G. Itani & H. Ueda (2010) Habitat use and horizontal distribution of the green mussel, Perna viridis, in Uranouchi Inlet, Kochi Prefecture. Sessile Organisms, 27: 41-50. (in Japanese)

Yamada, C., G. Itani & H. Asama (2009) Utilization of the non-indigenous green mussel, Perna viridis, by the native pinnotherid crab Arcotheres sinensis in Uranouchi Inlet, Kochi, Japan. Crustacean Research, 38: 70-76.

Jespersen, A., J. Lutzen & G. Itani (2009) Sperm structure and sperm transfer in Pseudopythina subsinuata (Bivalvia; Galeommatoidea). Zoologischer Anzeiger 248: 57-67.

Itani, G. (2008) Symbioses in estuarine burrows. Ishibashi, N. & Nawa, Y. (eds), "Parasitism and symbiosis", Tokai University Press, pp. 217-237. (in Japanese)

Nara, M., H. Akiyama & G. Itani (2008) Macrosymbiotic association of the myid bivalve Cryptomya with thalassinidean shrimps: examples from modern and Pleistocene tidal flats of Japan. Palaeogeography, Palaeoclimatology, Palaeoecology, 261: 100-104.

Yamada, C. & G. Itani (2008) The educational use of the "Yokonami Rinkai Experimental Station": lists of intertidal mollusks and beach shells. Bulletin of the Faculty of Education, Kochi University, 68: 165-170.

Itani, G. & T. Uchino (2005) A morphometric study of the burrowing mud shrimp Laomedia astacina (Decapoda: Thalassinidea: Laomediidae). Crustacean Research, 34: 87-94.

Kinoshita, K. & G. Itani (2005) Interspecific differences in the burrow morphology between the sympatric mud shrimps, Austinogebia narutensis and Upogebia issaeffi (Crustacea: Thalassinidea: Upogebiidae). Journal of the Marine Biological Association of the United Kingdom, 85: 943-947.

Itani, G., M. Izichi & H. Ueda (2005) Crab species collected from the burrows of Urechis unicinctus in Hiuchi-nada, the central Seto Inland SeaDCANCER, 14: 1-4. (in Japanese with English Abstract)

Itani, G. (2004) Distribution of intertidal upogebiid shrimp in Japan. Contributions from the Biological Laboratory, Kyoto University, 29: 383-399.

Itani, G. (2004) Host specialization in symbiotic animals associated with thalassinidean shrimps in Japan. Tamaki, A. (ed), Proceedings of the symposium on "Ecology of large bioturbators in tidal flats and shallow sublittoral sediments -from individual behavior to their role as ecosystem engineers." Nagasaki University, pp. 33-43.

Itani, G. (2004) Bopyrids | life history and species diversity. Nagasawa, K. (ed),"Aquaparasitology in the field in Japan", Tokai University Press, pp. 13-26. (in Japanese)

Itani, G. (2003) Symbiotic relationships in shrimp burrows. Asakura, A. (ed), "Biology of crustaceans | recent topics", Tokai University Press, pp. 233-253. (in Japanese)

Itani, G. & T. Uchino (2003) Burrow morphology of the goby Taenioides cirratus. Journal of the Marine Biological Association of the United Kingdom, 83: 881-882.

Itani, G. & M. Kato (2002) Cryptomya (Venatomya) truncata (Bivalvia: Myidae): association with thalassinidean shrimp burrows and morphometric variation in Japanese waters. Venus, 61: 193-202.

Itani, G. (2002) Two types of symbioses between grapsid crabs and a host thalassinidean shrimp. Publications of the Seto Marine Biological Laboratoy, 39: 129-137.

Itani, G., P. J. F. Davie & M. Takeda (2002) Taxonomic notes on Acmaeopleura balssi Shen, 1932 and A. toriumii Takeda, 1974 (Crustacea, Brachyura, Grapsidae) from Japanese waters. Bulletin of National Science Museum, Tokyo, Ser. A, Zoology, 28: 43-50.

Itani, G., M. Kato & Y. Shirayama (2002) Behaviour of the shrimp ectosymbionts, Peregrinamor ohshimai (Mollusca: Bivalvia) and Phyllodurus sp. (Crustacea: Isopoda), through host ecdyses. Journal of the Marine Biological Association of the United Kingdom, 82: 69-78.

Itani, G. (2001) Population characteristics of Upogebia yokoyai (Crustacea, Decapoda, Thalassinidea) at a tidal flat in southern Wakayama, Japan. Nankiseibutu, 43: 1-5. (in Japanese with English Abstract)

Itani, G. & H. Fujihara (2001) Carrying behaviour in a dorippid crab Heikea japonica (Arthropoda, Decapoda) - utilization of the sea pen (Cnidaria, Pennatulacea). Nankiseibutu, 43: 47-49. (in Japanese with English Abstract)

Itani, G. (2001) Shell allometry of Anomalocardia squamosa (Bivalvia: Veneridae) in Tanabe Bay. Nankiseibutu, 43: 161-162. (in Japanese with English Abstract)

Sato, M., H. Uchida, G. Itani & H. Yamashita (2001) Taxonomy and life history of the scale worm Hesperonoe hwanghaiensis (Polychaeta: Polynoidae) newly recorded in Japan, with special reference to commensalism to a burrowing shrimp Upogebia major. Zoological Science, 18: 981-991.

Saito, N., G. Itani & N. Nunomura (2000) A preliminary check list of Isopod Crustaceans in Japan. Bulletin of the Toyama Science Museum, 23: 11-107. (in Japanese with English Abstract)

Itani, G. (2000) Distribution of Acmaeopleura parvula (Brachyura, Grapsidae) in Izu-Ogasawara Archipelago. Nankiseibutu, 42: 69-71. (in Japanese with English Abstract)

Kato, M. & G. Itani (2000) Peregrinamor gastrochaenans (Bivalvia: Mollusca), a new species symbiotic with the thalassinidean shrimp Upogebia carinicauda (Decapoda: Crustacea). Species Diversity, 5: 309-316.

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